Read Ebook: The Social Direction of Evolution: An Outline of the Science of Eugenics by Kellicott William E William Erskine

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Let us have a concrete illustration. One of the simplest cases is that of the heredity of color in the Andalusian fowl which has been so clearly described by Bateson. There are two established color varieties of this fowl, one with a great deal of black and one that is white with some black markings or "splashes"; for convenience we may refer to these as the black and white varieties respectively. Each of these breeds true by itself. Black mated with black produce none but black offspring, white mated with white produce none but white offspring. Crossing black and white, however, results in the production of fowls with a sort of grayish color, called "blue" by the fancier, though in reality it is a fine mixture of black and white. At first sight we seem to have a gray hybrid race through the mixture of the black and the white races. Not so: for if we continue to breed successive generations from these blue hybrid fowls we get three differently colored forms. Some will be blue like the parents, some black like one grandparent, some white like the other grandparent. Not only this but we get certain definite proportions among these three classes of descendants. Of the total number of the immediate offspring of the hybrid blues, approximately one half will be blue like the parents, approximately one fourth black, and one fourth white like each of the grandparents. Now comes the most important fact of all. These blacks, bred together produce only blacks, the whites similarly produce only whites; the blues, on the other hand, when bred together produce progeny sorting into the same original classes and in the same proportions as were produced by the blues of the original hybrid generation. Their blacks and whites each breed true, their blues repeat the history of the preceding blues. No race of the hybrid character can be established: blues always produce blacks and whites, as well as blues. A summary of this history in graphic and diagrammatic form is given in Fig. 7.

This law of heredity was first discovered about forty-five years ago by Gregor Mendel, working with peas in the garden of the Augustinian monastery in Br?nn, Austria. His work curiously failed to arouse the interest of contemporary scientists and his results were soon completely lost sight of. The independent rediscovery of Mendel's formulas of heredity, about ten years ago, was probably the most important event in the history of biology and evolution since the publication of "The Origin of Species."

In the crossing of the black and white Andalusian fowls described above the phenomenon of dominance does not appear; when the two color characters are brought into a single individual neither appears alone, neither overcomes nor is overcome by the other. In the crossing of the black and white guinea pigs dominance is complete; when the two color characters are brought into a single individual only one color appears, the second becomes recessive, that is, it remains present as we know from the later history of such hybrids, but it is not visibly indicated. Besides the Andalusian fowls there are known several other instances of the absence of dominance and there are many cases where dominance is incomplete, i. e., where one character merely tends to dominate the other. And in a few instances dominance is irregular, i. e., sometimes one character dominates, at other times or under other circumstances it does not, as with certain forms of the comb or the feathering of the legs in the common fowl, or with the presence of an extra toe in the domestic cat, the rabbit, and guinea pig. And even in those cases where dominance is said to be complete the trained eye of the breeder can frequently distinguish between the hybrid and the pure bred dominant individuals. The phenomenon of dominance, therefore, is not an essential of the Mendelian theory although it is a frequent, we may say usual, relation.

That is, one fourth are pure black , one fourth pure white , and the remaining half are hybrids, black and white . The pure blacks again form germ cells, all possessing the determiner for blackness; the pure whites form germ cells all lacking the determiner for blackness; the hybrid blues produce again equal numbers of germ cells possessing and lacking the determiner for blackness. The relation of the germ cells and the organisms forming them and developing from them is shown in the diagram in Fig. 9.

In the more common cases where the phenomenon of dominance appears, as in the guinea pig, this is explained by saying that here a single determiner for blackness is somehow sufficient to produce the color. In such cases the black color observed may result either from a single or from a double black determiner in the germ which forms the organism. Only when the black determiner is entirely absent does the white color appear in the developed organism and the individual is then said to exhibit the recessive characteristic.

Another possible type of mating is that between a member of a pure race, either dominant or recessive, and a hybrid individual. This form of mating is very common in some of the pedigrees that we shall examine later. The results of such a mating, first between a hybrid and a recessive individual can be most easily described by considering a cross between black and white forms and expressing the result algebraically.

That is, returning to the example of the Andalusian fowls, the progeny will be one half hybrid blues and one half whites--no black at all. If the cross had been between black hybrid guinea pigs and white recessive specimens the result would have been half hybrid blacks and half pure whites.

Or supposing the mating to have occurred between the pure dominant and the hybrid the result would have been, in the fowls half pure black and half hybrid blue; in the guinea pig all the progeny would have been black, half pure blacks and half hybrid blacks.

This conception of the unit character is an extremely important element in the whole Mendelian theory and it has extended beyond the field of heredity and led to a radical change in our notions of what an organism really is. It is, of course, true in a sense that an organism is a unit, an organism is one thing; but at the same time it is true that an organism is fundamentally a collection of units, of structural and functional characteristics which are really separable things. A few of these units were mentioned in the first pages of this chapter and others are mentioned on a later page. They serve as the building blocks of organisms: individuals of the same species may be made up of similar combinations or of different combinations. One unit or a group of units may be taken out and replaced by others.

We shall have occasion in the next chapter to enumerate some of the human unit characters whose heredity has been traced and which have been found to Mendelize, but we may mention here a few Mendelizing units in other organisms in order to give some idea of the kind of character which behaves as a unit and of the range of the forms which have been found to show Mendelian phenomena in their heredity. Among the higher animals one might mention the absence of horns in cattle and sheep; the "waltzing" habit of mice and the pacing gait of the horse; length of hair and smoothness of coat in the rabbit and guinea pig; presence of an extra toe in the cat, guinea pig, rabbit, fowl; length of tail in the cat; and in the common fowl such characters as the shape and size of the comb, presence of a crest or a "muff," a high nostril, rumplessness, feathering of the legs, "frizzling" of the feathers, certain characters of the voice, and a tendency to brood. Among plants may be mentioned such characters as dwarfness in garden peas, sweet peas, and some kinds of beans; smoothness or prickliness of stem in the jimson weed and crowfoot; leaf characters in a great variety of plants; in the cotton plant a half dozen characters have been found to Mendelize; seed characters such as form and amount of starch, sugar, or gluten; flat or hooded standard in the sweet pea; annual or biennial habit in the henbane; susceptibility to a rust disease in wheat. We should not fail to mention that scores of color characters are known to Mendelize, such as hair or coat color and eye color in animals and the colors of flowers, stems, seeds, seed-coats, etc., in plants. The list of Mendelizing traits in different organisms now extends into the hundreds and is increasing almost weekly.

Before leaving the subject of Mendelism we should say that the phenomena, as described above in the Andalusian fowl and guinea pig, are among the simplest known. And while such simple formulas serve to describe the phenomena of heredity in a large number of instances, yet in a great many other cases the descriptive formulas are more complicated. We cannot in this place describe any of these complications. For a full discussion of these and of the whole subject of Mendelism the interested reader is referred to Professor Bateson's work on "Mendel's Principles of Heredity" . It must suffice to say here that in color heredity, for example, such ratios as 9:3:4 or 12:3:1 in the second filial generation instead of the more frequent 1:2:1 or 3:1 are explainable upon essentially the same relations as these simpler and more typical ratios. And further, many less usual Mendelian phenomena, which we cannot undertake to describe here, are associated with what the specialist technically terms "sex limitation," "gametic coupling," and the like.

It is often said that the Mendelian formula has a very limited applicability to human heredity. This is probably true if we consider carefully the grammatical tense in which this statement is made. And yet it is almost certainly true that heredity in man is to be described by this law. This apparent paradox is easily explained. The only characters whose history in heredity follows this formula are the unit characters. A complex trait is not heritable, as a whole, but its components behave in heredity as the separate units. It is perfectly well known that we are deeply ignorant regarding this phase of human structure. Our ignorance here is not the necessary kind, however, it is merely due to the newness of the subject--we have not had time to find out. How can we say that a complex trait is or is not inherited according to some form of Mendel's law when we do not know the nature of the units of which it is composed? We can make no statements about the Mendelian inheritance of such a trait until it is factored into its units. A considerable number of human characteristics are really known to be heritable according to this formula, enough so that several general rules of human heredity have been formulated. But it is also quite within the range of possibility that some traits really do not follow this law, although it cannot yet be said definitely that this is or is not the case. On the whole, then, we cannot, for the next few years, expect too much from the application of Mendel's laws to human heredity, however much this is to be regretted.

The method is the same as that employed by the statistician in measuring the relatedness of any two series of varying phenomena. If two quantities or characteristics are so related that fluctuations in the one are accompanied in a regular manner by fluctuations in the other, the two quantities or characters are said to be correlated. For instance, the temperature and the rate of growth of sprouting beans are related in such a way that increase in the former is accompanied in a regular way by increase in the latter; or the width and height of the head, or the total stature and the length of the femur similarly vary regularly together so that they are said to be correlated to a certain extent which can be measured. This correlation may result from the fact that one condition is a cause, either direct or indirect, of the other; or there may be no such causal relation between the two phenomena, both resulting more or less independently from a common antecedent condition or cause.

This phenomenon of correlation is not limited among organisms to the comparison of two or more different characters in a single series of individuals; it is applicable also to the comparison of two series of individuals with respect to the same characteristic. Thus we may compare the stature of a series of fathers with the same measurement in their sons. It is this form of correlation with which we are particularly to deal here. While it is not necessary to understand just how this subject is dealt with by the statistician we should know one or two of the elementary principles involved, in order to appreciate the statistical form of many statements about heredity.

The fact of regression is of considerable importance for the theory of evolution as well as for the subject of Eugenics when describing the phenomena of heredity in this statistical manner in whole groups without paying attention to particular individuals. Regression is found in all characteristics observed in this way, psychic as well as purely physical. "The father with a great excess of the character contributes sons with an excess, but a less excess of it; the father with a great defect of the character contributes sons with a defect, but less defect of it."

One further point remains to be considered, which applies not so much to coefficients of heredity as to coefficients of correlation in general, i. e., to the relatedness of two different characters or series of events in a single group of cases or individuals. This is that coefficients of correlation may be either positive or negative. That is, the real limits of the value of the coefficient are plus one and minus one. The example given above of stature of fathers and sons gives a positive coefficient. Whenever the deviation from the average of one group is accompanied in the second group by a deviation in the same direction, the coefficient is positive. A negative correlation means that deviation from the average in a given direction in the first group is accompanied in the second group by a deviation in the opposite direction. If we imagine that as one measurement increased above its average a second related measurement decreased below its average the correlation in such a case would be negative. For instance, if we measured the relation between the number of berry pickers employed and the quantity of berries remaining unpicked, in a number of different fields we would get a negative correlation coefficient. Some organisms are formed in such a way that increase in one dimension, such as length, is associated with decrease in another, such as breadth; measurement of the relatedness of these dimensions would give a coefficient of correlation that might be very high, indicating a considerable relation in the deviations, but it would be negative. In an instance of negative correlation the relation is that of "the more the fewer." As we shall see presently, a negative correlation may be just as important and significant as a positive correlation.

The application of the principles of heredity to our subject of Eugenics is of such great importance that it is reserved for separate consideration in the next chapter. We may, therefore, devote the remainder of this chapter to the consideration of data of another kind, which are commonly treated by this same method of determining correlation coefficients between two sets of varying phenomena in order to determine whether there is any actual relation between them or not. This will serve to illustrate the use of this method.

We shall turn then to the subject of differential or selective fertility in human beings and consider its relation to Eugenics. As a starting point we may take the self-evident statement that a group of organisms will tend to maintain constant characteristics through successive generations only when all parts of the group are equally fertile. If exceptional fertility is associated with the presence or absence of any characteristic the number of individuals with or without that trait will either increase or diminish in successive generations, and the character of the distribution of the group as a whole will gradually become altered, the average moving in the direction of the more fertile group. Or if infertility is so associated, then the average of the whole group moves away from that condition. Eugenically, then, we should ask whether in human society there is at present any such association of superfertility or infertility with desirable or undesirable traits. It is obviously the aim of Eugenics to bring about an association of a high degree of fertility with desirable traits and a low degree of fertility with undesirable characteristics.

First, let us look at certain data gathered relative to the size of the family in both normal and pathological stocks . In order that a stock or family should just maintain its numbers undiminished through successive generations and under average conditions, at least four children should be born to each marriage that has any children at all.

TABLE II

Deaf-mutes, England Schuster Probably complete 6.2 Deaf-mutes, America Schuster Probably complete 6.1 Tuberculous stock Pearson Probably complete 5.7 Albinotic stock Pearson Probably complete 5.9 Insane stock Heron Probably complete 6.0 Edinburgh degenerates Eugenics Lab Incomplete 6.1 London mentally defective Eugenics Lab Incomplete 7.0 Manchester mentally defective Eugenics Lab Incomplete 6.3 Criminals Goring Completed 6.6 English middle class Pearson 15 years at least, begun before 35 6.4 Family records--normals Pearson Completed 5.3 English intellectual class Pearson Completed 4.7 Working class N.S.W. Powys Completed 5.3 Danish professional class Westergaard 15 years at least 5.2 Danish working class Westergaard 25 years at least 5.3 Edinburgh normal artisan Eugenics Lab Incomplete 5.9 London normal artisan Eugenics Lab Incomplete 5.1 American graduates Harvard Completed 2.0 English intellectuals Webb Said to be complete 1.5

All childless marriages are excluded except in the last two cases. Inclusion of such marriages usually reduces the average by 0.5 to 1.0 child.

The table given shows clearly what stocks are maintaining, what increasing, and what diminishing their numbers.

CORRELATION COEFFICIENT. With number of males engaged in professions -.78 With female domestics per 100 females -.80 With female domestics per 100 families -.76 With general laborers per 1,000 males +.52 With pawnbrokers and general dealers per 1,000 males +.62 With children employed, ages 10 to 14 +.66 With persons living more than two in a room +.70 With infants under one year dying per 1,000 births +.50 With deaths from pulmonary tuberculosis per 100,000 inhabitants +.59 With total number of paupers per 1,000 inhabitants +.20 With number of lunatic paupers per 1,000 inhabitants +.34

But the sign changes and becomes positive when we come to other comparisons. When we count the relative number of pawnbrokers and general dealers, of "general laborers" , of employed children between the ages of ten and fourteen, of persons living more than two in a room, when we consider the infant death rate, the death rate from pulmonary tuberculosis, and the relative number of paupers,--then we find the signs of the coefficients are all positive, and on the average the coefficients are more than 0.50--a moderate to high degree of regularity of the relation. The districts characterized by the larger numbers of such individuals or by higher death rates of these kinds, are at the same time the districts where the birth rates are the higher.

Sidney Webb has recently published an account of the birth-rate investigations undertaken by the Fabian Society with a view to determine the causes leading to the rapidly falling birth rate in England. During the decade previous to 1901 the number of children in London actually diminished by about 5,000, while the total population increased by about 300,000. As far as they bear upon this phase of the subject his results fully confirm these we have been considering. The falling off is chiefly in the upper and middle classes, in the classes of thrift and independence, and it has occurred chiefly during the last fifty years. Webb cannot find that this is due to any physical deterioration in these classes; it is due to a conscious and deliberate limitation of the size of the family for what are thought prudential and economic reasons.

An actual reduction in the number of children may not be an unmixed evil. A falling birth rate may be a good sign. This is partly a question for the political economist. "Suicide" may be a socially fortunate end for some strains. But when, in either a rising or a falling birth rate, we find a differential or selective relation, then the subject is eugenic. If the higher birth rate is among the socially valuable elements of each different class the Eugenist can only approve; to bring about such a relation is one of his aims. What we really find, however, is the undesirable elements increasing with the greatest rapidity, the better elements not even holding their own.

We have here a result that has very important bearings upon the value to the race of the large family and of the danger of the small family. The small family of one, two, or three children contributes on the average much more than its share of pathological and defective persons. No matter just now what the causes are, they seem to be more or less beyond remedy. The result for the future, however, must be reckoned with. This relation has important bearings upon the custom of primogeniture as well as upon the eugenic values of the large family.

The subject before us illustrates the direct bearing of science upon moral conduct and upon statecraft. The scientific study of man is not merely a passive intellectual viewing of nature. It teaches us the art of living, of building up stable and dominant nations, and it is of no greater importance for the scientist in his laboratory, than for the statesman in council and the philanthropist in society.

HUMAN HEREDITY AND THE EUGENIC PROGRAM

HUMAN HEREDITY AND THE EUGENIC PROGRAM

"A breed whose proof is in time and deeds; What we are, we are--nativity is answer enough to objections."

A few years ago official recognition was taken of the disturbing fact that the annual wheat yield of Great Britain was grossly deficient in both quantity and quality. In 1900 The National Association of British and Irish Millers, with almost unprecedented sagacity, raised a fund to provide for a series of experiments under the direction of a competent biologist, in order to discover if possible some means of restoring the former yield and quality of the native wheats. The story of the result reads like a romance. The experimenter--Prof. R. H. Biffen--collected many different varieties of wheat, native and foreign, each of which had some desirable qualities, and studied their mode of inheritance. Now, after only a few years of experimentation a wheat has been produced and is being grown upon a large scale in which have been united this desirable character of one variety, that character of another. From each variety has been taken some valuable trait, and these have all been combined into one variety possessing the characteristics of a short full head, beardlessness, high gluten content, immunity to the devastating rust, a strong supporting straw, and a high yield per acre. A wheat made to order and fulfilling the "details and specifications" of the growers.

Manitoba and British Columbia opened up whole new lands of the finest wheat-growing capacity, but the season there is too short for the ripening of what were the finest varieties. This new specification was promptly met and the early ripening quality of some inferior variety was transferred to the varieties showing other highly desirable qualities, and these countries are now producing enormous quantities of the finest wheat in the world.

All of this has been made possible by the discovery, mentioned in the preceding chapter, that many characteristics of organisms are units and behave as such in heredity; they can be added to races or subtracted from them almost at will. Pure varieties breeding true can be established permanently by taking into account the Mendelian laws of heredity. Similar results have been accomplished in many other plants and in many animals. A cotton has been produced which combines early growth, by which it escapes the ravages of the boll weevil, with the long fiber of the finest Sea Island varieties. Corn of almost any desired percentage of sugar or starch, within limits, can be produced to order in a few seasons. The hornless character of certain varieties of cattle can be transferred to any chosen breed. Sheep have been produced combining the excellent mutton qualities of one breed with the hornlessness of another, and with the fine wool qualities of still a third. And so on from canary birds to draft horses. New races can be built up to meet almost any demand, with almost any desired combination of known characters, and these races remain stable. Possibilities in this direction seem to be limited only by our present and rapidly lessening ignorance of the facts of Mendelian heredity in organisms--facts to be had for the looking.

What is man that we should not be mindful of him? Why should we utilize all this new knowledge, all these immense possibilities of control and of creation, only for our pigs and cabbages? In this era of conservation should not our profoundest concern be the conservation of human protoplasm? "The State has no material resources at all comparable with its citizens, and no hope of perpetuity except in the intelligence and integrity of its people." As Saleeby puts it: "There is no wealth but life; and if the inherent quality of life fails, neither battle-ships, nor libraries, nor symphonies, nor Free Trade, nor Tariff Reform, nor anything else will save a nation."

In this work of the creation and establishment of new and valuable varieties, two essential biological facts are made use of. The raw materials are furnished by variation--by the fact that there are individual and racial differences. The means of accomplishing results are furnished by heredity--the fact that offspring resemble the parents, not only in generalities, but even in particulars, and according to certain definite formulas.

At the outset we should say that the knowledge of human heredity is as yet largely of the statistical sort. We know how a great many characters are inherited, on the average. The subject of Mendelian heredity is so new that there has been hardly time to investigate more than a few human characteristics from this point of view. Certain conditions add to the difficulties here. First, many, probably most, of the more important human traits are complexes, not units, and it is a long and difficult process to analyze them into their units, with which alone Mendelism deals. Second, in human society we cannot carry on definite experiments under controlled conditions, directed toward the solution of some concrete problem in heredity. It is true that Nature herself is making such experiments constantly, but at random, and rarely under ideal conditions of what the experimenter calls control or check. We have first to seek and find them out, and when they are found we often discover that there are lacking many of the facts essential to a complete or satisfactory analysis of the facts displayed. The comparatively small size of the human family sometimes makes it difficult to get data sufficiently extensive to be really significant. And the long period that elapses between successive human generations adds to the difficulty of getting precise information, for in dealing with the heredity of some traits comparisons must be made with individuals of the same ages, and the period of observation of a single observer seldom exceeds the duration of a single generation. Yet in spite of all these difficulties we have a fairly broad and exact knowledge of human heredity in respect to some characteristics.

Human heredity involves both physical and psychical characters--both the body and the mind are concerned. Among other animals little if anything is known regarding psychic inheritance, but the physical traits of men are inherited in just the same ways and to the same degrees as in animals. This degree or intensity of inheritance may be expressed in coefficients of heredity between the groups of relatives being compared. To mention a few examples of coefficients for physical traits we have the following:

CHARACTER OBSERVED PARENTAL FRATERNAL COEFFICIENT COEFFICIENT Stature .49-.51 } .51-.55 } Span .45 } .55 } Fore Arm .42 } .47 .49 } .53 Eye Color .55 } .52 } Hair Color .57 - Average Hair Curliness .52 Head Measurements-three .55 - " Cephalic Index .49

We might give many others, but it is unnecessary. Notice that these parental and fraternal coefficients group about an average value of about .50 or slightly less. Similar coefficients have been worked out for other degrees of relationship; thus grandparental coefficients are about .25.

Stated briefly, in less exact terms, these coefficients mean that, with respect to such traits as deviate from the group average, the resemblance of brothers and sisters to each other or of children to their parents is, on the whole, approximately mid-way between being complete in its deviation from the average and in not deviating at all from the average in the direction of the fraternal or parental characteristic. Grandchildren tend to deviate from the group average only about one fourth as far as their grandparents. It should be remembered that these are statistical and not individual statements, and that as many "exceptions" will be found in the direction of greater resemblance as in that of lesser resemblance.

One of the present objects of the student of heredity, perhaps his chief object, is to be able to state the facts of human heredity in Mendelian terms, reducing many of the complex human traits to their simpler elements. Some of the chief objections to the use of the statistical formula of heredity are that apparently it is applicable only to the fluctuating variabilities of organisms; that it rarely takes into account the presence of true variations or mutations--and we have seen that it is just these characters that are of the greatest value in evolution; and that heredity is after all fundamentally an individual relation which loses much of its definiteness and significance when we merge the individual in with a crowd. To some these seem fatal objections to any use of the statistical formula and it is certainly true that they greatly limit its value. But for the present at least the statistical statement of certain facts of heredity is still useful in this bio-social field. We may therefore use the statistical formulas of heredity as a kind of temporary expedient, enabling us to make statements regarding inheritance of certain characters in the group or class, pending the time when we shall be able to give the facts a more precise and more "final" expression in Mendelian formulas. Many human traits are indeed already known to Mendelize. Most of these are, however, "abnormal" traits or pathological conditions; we are still in the dark regarding the actually Mendelian or non-Mendelian inheritance of most of man's normal characteristics. We might enumerate the following Mendelizing human characters--eye color, color blindness, hair color and curliness, albinism , brachydactylism , syndactylism , polydactylism , keratosis , haemophilia , nightblindness , certain forms of deaf mutism and cataract, imbecility, Huntington's chorea .

Turning now to the inheritance of mental traits and including, of course, moral traits here as well, we find that we are almost entirely limited to the statistical statement of results. Pearson found upon examining data from a large number of school children, brothers and sisters, that the coefficients of heredity between them were the same as for their physical traits. His results are summarized in Figure 12. The physical traits measured were, in the order plotted in the figure--health, eye color, hair color, hair curliness, cephalic index , head length, head breadth, head height. These gave an average of .54 in brothers, .53 in sisters, and .51 in brothers and sisters. The psychical traits in order were--vivacity, assertiveness, introspection, popularity, conscientiousness, temper, ability, handwriting. The corresponding averages were .52, .51, .52.

Galton's pioneer works on "Hereditary Genius," "English Men of Science," and "Natural Inheritance" showed with great clearness the fact of mental and moral heredity. Wood's recent extensive study of "Mental and Moral Heredity in Royalty" shows the same thing, although not all the results of these investigations are given in mathematical form. Little can be said regarding Mendelian heredity of mental traits because the psychologist has not yet told us how to analyze even the common and simpler psychic characters into their fundamental units; since we do not know what the mental hereditary units are, obviously we cannot work with them. Much of our knowledge in this field does not permit of very accurate summary, though pointing indisputably to the fact of mental inheritance in spite of the very great influences of training and education, environment and tradition, in moulding the mental and moral characteristics--influences with much greater effect here than in connection with physical characters.

Galton studied the parentage of 207 Fellows of the Royal Society, a Fellowship which is a real mark of distinction. He assumed that one per cent of the individuals represented by the class from which his observations were drawn, that is the higher intellectual classes, might be expected to be "noteworthy": among the general population the average is really about one in 4,000 or one fortieth of one per cent. On the one per cent basis Galton found that Fellows of the Royal Society had noteworthy fathers with 24 times the frequency to be expected in the absence of heredity; noteworthy brothers with 31 times the expected frequency; noteworthy grandfathers 12 times; and so on through various grades of relationship.

Schuster examined the class lists of Oxford covering a period of 92 years and found that first honor men had 36 per cent first or second honor fathers; second honor men had 32 per cent first or second honor fathers; ordinary degree men 14 per cent first or second honor fathers. These percentages are far in excess of that to be expected--perhaps 0.5 per cent--on the assumption that ability is not inherited. Schuster also determined the coefficients of heredity between fathers and sons as regards intellectual ability, the evidence being class marks in Oxford and Harrow; these he found to be about .3 for the parental relation and .4 for the fraternal. The intensity of heredity in many forms of insanity has been determined and this runs up much higher--.57 parental and .50 fraternal.

It is clear I take it, that the fact of human heredity does not concern only physical traits but extends to psychical traits as well, and with about the same intensity. This fact has been found true also for still less analyzable characters such as length of life, fertility or infertility and the like, and again about the same intensity of resemblance is found.

Human heredity is a fact then just as human variability is a fact. We have truly the raw materials and the means for racial improvement. The ability to direct the evolution of the human race makes this our supremest duty.

The defect we have just been considering is dominant. Many defects are recessive, i. e., transmitted though not exhibited by a hybrid individual. Viewed from the standpoint of the character of the offspring, mating with such a person would be unfit only when both persons were similarly recessives. Such a chance similarity would be likely only in cases of blood relationship. Here lies the scientific basis for many of the legal restrictions against cousin marriage or the marriage of closer relatives, for here, although both persons may appear normal, the chances for latent ills appearing in the progeny in a pure and permanently fixed condition are greatly increased. Of course the same relation holds for characteristics which are not defects but really valuable traits. Marriage of cousins possessing valuable characters, whether apparent or not, might be allowed or encouraged as a means of rendering permanent a rare and valuable family trait which might otherwise be much less likely to become an established characteristic. Some discrimination should be exercised in the control, legal or otherwise, of such marriages.

Fig. 14 gives a brief pedigree of a family in which polydactylism occurs. This is a condition in which one or more additional or supernumerary fingers or toes are present in the extremities. The Mendelian character of the heredity of this defect is less clear than in the preceding, yet there are many indications that this is really an illustration of a complex Mendelian formula. Probably if the parentage of the individuals marrying into this family were known we should be able to give a complete formula. At any rate the pedigree illustrates the unfit character of the matings with affected persons, for in no instance has such a marriage resulted in the production of fewer than one half affected offspring.

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