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Transcriber's note: A few typographical errors have been corrected: they are listed at the end of the text.

CONCHOLOGICAL MANUAL.

G. B. SOWERBY, JUN.

SECOND EDITION.

CONSIDERABLY ENLARGED AND IMPROVED.

LONDON:

HENRY G. BOHN, YORK STREET, COVENT GARDEN.

PREFACE TO THE FIRST EDITION.

It may be necessary in introducing this little volume, to state, that it is strictly conchological, and that it is compiled for the use not only of those who wish to acquire an elementary acquaintance with the subject, but also of authors and others, who, desirous of extending their knowledge and pursuing their researches, require a book of reference, containing a general outline of what has been done by those who have trodden the same path before them. It has been thought advisable, for general convenience, to arrange the principal part of the information in alphabetical order: adding tables of the systems of Lamarck and De Blainville, to facilitate the systematic pursuit of the science.

Persons of the class first alluded to, will find great assistance in the explanation of technical words, their application being further illustrated, in most cases, by a reference to the figures; and, although they might have been multiplied, it is trusted that enough are given for every useful purpose.

The definition of the Classes, Orders, Families, and Genera, in the system of De Blainville, and a tabular view, are presented for the use of those who prefer it, or who wish to compare it with that of Lamarck.

In the explanation of the figures, will be found a systematic arrangement of shells, according to Lamarck, including the names of genera established or proposed since the publication of his system. The descriptions of established genera have been rendered as concise and clear as possible. It is hoped that no essential characters are omitted, and that those living authors, whose proposed generic distinctions have been passed over in a few words, will not have to complain of want of justice in the attempt to interpret their meaning.

In most cases the generic name will be found accompanied by its derivation. This has been done, in the hope of assisting the memory by associating the meaning of a term with some peculiarity in the thing described. At the end of each description of a genus, some general observations occur, pointing out the principal character which distinguishes it from others, to which it is nearly allied; and also stating the geographical or geological distribution and habits of the animal.

The above descriptions and definitions are illustrated by a series of plates, containing above 500 etchings of nearly as many proposed or established genera, arranged in Lamarckian order, so as to show at a glance all the generic forms of each family. And, although from their number, they could not be very highly finished, it is hoped that they will be found characteristic.

The compiler cannot replace his pen without acknowledging, with filial gratitude, the kind assistance of one who has sacrificed much of his time in bringing his knowledge and experience to bear upon the correctness and utility of this humble attempt to remove some of the difficulties to which the commencement of this, as well as of every other study, is exposed.

PREFACE TO THE SECOND EDITION

The favourable reception and rapid sale of the first edition of the Conchological Manual having rendered a second necessary, the Author takes this opportunity of explaining the nature of the alterations which have been made. In doing this, he has to thank his friends for their suggestions, which, together with his own increased knowledge and experience, enable him to present a more complete and satisfactory work to the scientific public.

For the further convenience of those who are studying the rudiments of the science, an entirely new Introduction is given, in which, commencing with the structure and gradual developement of the shell, the author has endeavoured to explain the general principles of Conchology in systematical order. This Introduction is illustrated by 100 wood-cuts, which will be found greatly to assist the Student.

The definitions have been rendered more full and complete than before, and the Author has profited by some manuscript notes communicated by a scientific friend, to whom he desires to present his humble acknowledgments. Upwards of four hundred explanations have been given of words which did not appear in the former edition, three-fourths of which are of generic and subgeneric names.

A large number of notes, referring to the geographical distribution of the genera, have been added from the pen of Mr. G. B. Sowerby, Senior.

The plates have been carefully improved; and three, containing upwards of eighty figures, have been added.

On the whole, it will be found that the amount of matter has been nearly doubled; all the defects, as far as they have been discovered, have been removed, and every means used of making the present edition as useful as possible.

INTRODUCTION.

The Science of Conchology affords a very delightful and instructive amusement for the leisure hours of those who, retiring occasionally from the gaieties of fashionable life, seek pleasure in the quiet contemplation of some of the smaller, but not less wonderful operations of creative wisdom. And, although the study of shells would be more complete, and rank higher in the scale of philosophical pursuits, were it always accompanied by that of the animal inhabiting them, it nevertheless presents means of intellectual gratification, to many who cannot follow it beyond the cabinet and the boudoir. These may examine with admiration and mental improvement, the beautiful colouring and architecture of these wonders of the deep, they may exercise their taste and judgment in the selection and arrangement of specimens, and their discrimination in detecting and appreciating the distinctions upon which the arrangement is founded.

It is but little that can be known of the subject without forming a collection of greater or less extent; for, as it would be uninstructive merely to delight the eye with the bright colours and elegant form of shells, without possessing correct information respecting them, so it would be insipid and useless to learn technicalities without being acquainted by personal observation with the subjects to which they are applied. The first endeavour should, therefore, be to obtain a few shells as examples of the larger divisions, and, when these are understood, to proceed with the smaller groups, until a collection be formed to represent as many generic forms as possible. It may be as well here to advise those who are forming a collection to be very particular in every practicable instance to have the shells properly named at the time of purchasing; as it will save much trouble, and materially assist in the attainment of the desired object. To this end, recourse should be had to those naturalist tradesmen, who unite the attainment and diffusion of real scientific knowledge with their commercial pursuits.

Supposing, however, that the person who desires to learn the science, possesses a small parcel of unarranged and unnamed shells, without any previous acquaintance with the subject, the following introductory explanations, are drawn up with the view of enabling him, without further assistance, to obtain a general insight into its principles, equal to that of those who have studied it long and laboriously. To effect this, he must read them, carefully comparing the descriptions with the figures referred to, and with the specimens which he may have at command.

Let none be discouraged by the number of generic distinctions proposed and adopted in modern times; for if well defined, they will be found to facilitate rather than encumber the science. The knowledge of species must be the foundation of every system, and the greater their number, the more necessary it becomes to subdivide them; if, for instance, all the species now known were to have been included in the 50 genera of Linnaeus, a single genus would have contained many hundreds of incongruous species, in which case it would be much more difficult to remember them, than if they were to be divided into a far greater number of genera. Every well marked division, however arbitrary its limits, tends to simplify the subject, and to facilitate the researches of the student.

NATURE OF THE SCIENCE.

Conchology is the study of shells, viewed and described as to what they are either in themselves, or in relation to the soft, inarticulate animals which produce them, and of which they form a part. These animals are called MOLLUSCA, and perhaps the best general description of them will be found in De Blainville's "Manuel de Malacologie et de Conchyliologie." The following is a translation, "Animal in pairs, the body and its appendages soft, inarticulate , enveloped in a muscular skin, commonly called the mantle, which is extremely variable in form, and has developed either within or upon it a calcareous portion, consisting of one or several pieces, commonly called a SHELL."

The term Mollusca was formerly restricted to those soft animals which were destitute of shells, although possessing in other particulars, the characters described above, and it was used in order to distinguish them from the TESTACEA, which were covered or internally supported by calcareous parts. In the system of Linnaeus, the soft portions are first arranged under the general designation of "Vermes Mollusca," and described without regard to the presence, absence, or character of the shells; and then the shells are separately characterized under the appellation of "Vermes Testacea," without any further notice of the animal, than an indication of the genus to which it belongs; thus the animal of Cypraea is said to be a Limax, and that of Tellina a Tethys.

The nearest approach to correctness, and the most philosophical method of study will be found in the modern system, adopted by Lamarck and his followers, of observing these animals as a whole, and arranging them according to the assemblage of characters which they present; of course taking into consideration the existence or non-existence, form and structure of the shell, on the same principle, which, in arranging the vertebrated animals would lead us to study the hair, hoof, nails, claws, &c. as well as the other parts.

On account of these and other considerations, it has been thought advisable that the present undertaking should bear a purely conchological character. The peculiarities of the shells alone being detailed for the assistance of those who collect and study them, while at the same time, in deciding upon their affinities and places, in the arrangement, it will be necessary to take advantage of the conclusion to which those have arrived, who have studied the animal in all its parts. And the conviction must be expressed, that if ever a complete Natural System shall be formed it will result from the labours of the last mentioned class of naturalists.

DEFINITION OF A SHELL.

A true shell is composed of one or more calcareous pieces, commonly called valves, each piece formed by a series of layers, applied obliquely upon each other, in such a manner that each new layer begins within, and terminates a little in advance of the one before it.

STRUCTURE AND GROWTH.

We shall now endeavour to describe the manner in which the growth of each separate valve, or each regularly formed shell, proceeds from the nucleus.

In both cases, the nucleus is generally of a more horny and transparent composition than the parts subsequently produced. As soon as the animal is hatched, or, in other words, leaves the egg or body of the parent, of course it begins to increase in size, and to require a corresponding enlargement in the shell. To effect this, a small quantity of mucus substance, secreted by the mantle of the animal, is deposited on the edge of the aperture. When this is dry and become sufficiently hard, it is lined by a more calcareous secretion; and these together form a new layer, which is followed by others in succession; each new layer being larger than the one that preceded it until the whole being complete, the full-grown animal is invested with a shell commensurate with its own proportions. Thus from the apex or nucleus the formation proceeds, as it were, downwards, taking the shape of the part which secretes it, on which it is in a manner moulded.

The nucleus, or first formed portion, may for technical purposes be considered, mathematically, as the apex of a spiral cone. And here it must be observed, that whether the shell consist of one or several pieces, each piece has a separate nucleus, and the process of formation is separately repeated with each. The word cone is used for convenience, and its meaning extended so as to include all those structures which commencing at a point enlarge downwards.

CLASSIFICATION.

NUMBER OF PIECES, OR INDEPENDENTLY FORMED PARTS.

But although this arrangement may appear at first sight perfectly easy and plain, some explanation will be necessary in order to guard the student against understanding the above expressions in their strictest sense, without qualification. Thus the univalves are said to consist of a single piece, or spiral cone; but it would be more correct to speak of this piece as forming either the whole or the principal part of the shell: for there is in many instances, a much smaller flattened piece attached to the foot of the animal, which being drawn in when it retires, closes the aperture as with a kind of door, to which in fact the word valve might be very properly applied; it is called however the OPERCULUM, and the little horny plate, frequently drawn out by means of a pin from the aperture of a periwinkle, will present a familiar example.

Another distinction, leading to important results in classification, is that which is derived from the nature of the element breathed by the Mollusc. And although this consideration belongs more especially to the study of the animal itself, yet the habits of the animal materially influence the structure of the shell.

In the former, represented by the cut, fig. 8, the tendon passes through a perforation in the upper valve; and in the latter, Mytilus, fig. 9, the byssus passes out between the valves.

Before proceeding to explain the characters of the different groups, according to the modern system of classification, it may be desirable to explain the terms by which the different parts and characters are described, and to shew the manner in which the shells are measured. For this purpose we shall treat of the general divisions separately. We begin with

UNIVALVE SHELLS.

In considering Univalves merely with reference to their mathematical construction, the first point demanding our attention is, whether they are symmetrical or non-symmetrical, or, in other words, whether a straight line drawn through the shell would divide it into two equal parts. The greater part of univalves are non-symmetrical, being rolled obliquely on the axis; but many are symmetrical, being rolled horizontally on the axis. The Nautilus presents an illustration of the latter; the Snail is a familiar example of the former.

The next variation in symmetrical univalves is to be observed in the tubular, curved form, the example of which will be the Dentalium, fig. 12.

CHAMBERED SHELLS.

In the above section of a Nautilus, fig. 13, diminished in size, showing the whorls and chambers , it will be seen that the edges of the septa are formed in one simple curve. In fig. 14, the upper part of an Ammonite, the undulating line will be seen; and in fig. 15 a specimen is given of the arborescent septa.

NON-SYMMETRICAL UNIVALVES.

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