Read Ebook: Mémoires de Luther écrits par lui-même Tome II by Luther Martin Michelet Jules

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Definition--Variation of migration.

Direction of passage--The potentiality of flight--Habit of wandering--Memory--Extension of range--Influence of Temperature--Desire for Light--Glacial Epoch--Food Basis--Sexual Impulses--Competition.

Route or Broad Front--Coasting--Fly-lines--Isepipteses --Land-bridges--Coast Lights.

Altitude of Normal Migration--Variation in Speed--Effect of Wind.

Route Finding--Use of Memory--Eyesight--Errors--Guidance of Young--Beam Winds--Homing of Terns.

The Swallow--Variation in Distances--Marking Birds--Results--Routes of the Golden Plover--Evolution of the Routes.

Knowledge of Approaching Weather--Favourable and Unfavourable Conditions--Importance of Winds--Cyclonic and Anticyclonic Winds--Continental Migration.

Contrary Winds--Lighthouses and Lightships--Leeward Drift--Catastrophes.

Literature--Hibernation--Carriage of Small by Large Birds.

Trans-Atlantic Migration--Ship-borne Wanderers --Storm-blown Birds--Casual Wanderers--Swimming and Walking.

BIBLIOGRAPHY 131

INDEX 135

LIST OF MAPS

FACING PAGE

Map showing the range of the American Golden Plover, with its known migration route 76

Map showing the evolution of the migration route of the American Golden Plover 78

Map showing the evolution of the migration route of the Eastern or Pacific Golden Plover 80

Map to show that a bird leaving Norway, near Aalsund, might be carried round the British Islands in twenty-four hours. The arrows indicate the actual directions and force of wind at the times marked during a slow-travelling circular storm in autumn 1901. Speed of bird about twenty-five miles per hour 98

THE MIGRATION OF BIRDS

MIGRATION OF BIRDS

Migration is the act of changing an abode or resting place, the wandering or movement from one place to another, but technically the word is applied to the passage or movement of birds, fishes, insects and a few mammals between the localities inhabited at different periods of the year. The wandering of a nomadic tribe of men is migration; the mollusc, wandering from feeding ground to feeding ground in the bed of the ocean, migrates; the caterpillar migrates from branch to branch, even from leaf to leaf; the rat leaves the ship in which it has travelled and migrates to the granary; we pack our goods, hire a removing van and migrate to a new abode. The word migration thus applied may be literally correct but it fails to convey the generally accepted meaning, and the expression Bird Migration suggests periodical and regular movement, the passage as a rule between one country and another.

The popular application of a term does not do away with the need of definition, especially as there are many complicated phases of migration. The migration of birds is as a rule between the breeding area or home and the winter quarters, but there are many migrants which never reach breeding quarters in spring, and many others which leave the regular breeding quarters or the place of residence in winter to perform a very real migration under peculiar stress of circumstances. Again the spasmodic movements of certain gregarious species, which at irregular intervals change their location in large numbers to take up their abode in another part of the range, is really migration, though it is now usually described as irruption, incursion or invasion.

Newton says that bird migration is "most strangely and unaccountably confounded by many writers with the subject of Distribution," but the very act of the bird which extends its range, the first step in distribution, is migration. The histories of present-day distribution and migration are irrevocably interwoven; as Mr P.A. Taverner remarks , "migration is a dispersal, and conversely, this dispersal, as it manifests itself, is migration," whilst distribution is the outcome of dispersal.

Broadly speaking, all birds migrate, though the length of the journey varies in different species, and in some cases in individuals of the same or closely allied species, from the merest change of elevation to a voyage almost as wide as the world itself. The sedentary red grouse nests on the moors, often less than 1000 feet above the sea, but "when snow-bright the moor expands" it feeds and resides in the cultivated valley, and as shown by the committee appointed to study grouse disease, not infrequently migrates from range to range across wide valleys. Many tropical birds, usually considered non-migratory, are subject to short movements, the origin and purpose of which is search for food and safe nesting places.

The knot breeds in countless numbers in Arctic Greenland and America, so far north that only a handful of ornithologists have traced its home; it travels south in summer so far as Damara Land. The Arctic tern has a northern breeding range extending perhaps as far north as that of any bird, and it has been taken far to the south of South America in the Antarctic regions; if the thesis that the further north the bird goes in summer the further south it travels in winter is correct, as it can be proved to be with some species, some of these terns must annually travel about 22,000 miles . Between these extremes are an endless variety of distances travelled and methods of migration, with striking differences in the performances of individuals of the same species. Take one instance, a song thrush reared in a nest in our own garden. We may see and recognise this bird up to the middle of July, but what trained ornithologist can, yet, say with certainty where that bird will be by the end of the month or in three to four months time? We know that all through the winter there are some song thrushes near the house, and that they are the birds which not only begin to sing early but actually nest with us; we know too that before there is any marked immigration of northern thrushes there is a recorded emigration from our southern coasts, presumably of thrushes which have nested with us, beginning towards the end of July; further we know that there is an autumn immigration of Scandinavian or other northern song thrushes, sub-specifically distinct to the expert eye, and some, small and dark, whose origin is by no means proved, as well as later emigrations of birds to the Continent or Ireland, both regular and occasioned by exceptional weather. Will our young July thrush remain in England or will it join one of these streams, and if so which? We do not know yet. I repeat "yet," for the study of races, sub-species or local variations is commanding more and more attention; the patient work of the "splitters," scorned by the old school of "lumpers," will eventually solve many of the problems of to-day.

The ancients--a usefully ambiguous term--realised that birds migrated; our immediate forefathers of two or three centuries ago realised that certain birds vanished in winter and wondered how; and within modern times the phenomena of migration, the "mystery of mysteries," has been the subject of much study, speculation, and literary exposition. Indeed a full bibliography of migration would be a considerable volume. Even workers within the last few years have declared that certain phenomena were beyond human understanding, only to be explained by instinct, a word capable of most varied interpretation. In truth there is much to learn, much to which we must still answer--we do not know; but the speculative theory of yesterday is now either myth or fact, and the theory of to-day may be proved true and add something to the data of which knowledge is built. The wildest speculations, based on slender locally ascertained facts or on no foundation whatever except the fertility of the brain, have been offered as solutions of the mysteries; the literature of migration is a jumble of contradictions. John Legg, in 1780, said "In relating so many instances of unparalleled credulity, I confess I cannot suppress the irascible passion" , and Herr Otto Herman, only a few years ago, pointing out the ingenious dogmas "void of every firm foundation," says that "really it is a field in which every thinking ornithologist may create new theses to any extent and more or less incredible" .

Herr Herman's system of "ornithophaenology," the accumulation of substantiated observations and facts, will not prove everything, but his work in Hungary, that of Dr Merriam and Mr Cooke in America, and of Mr W. Eagle Clarke in Britain, each aided by a numerous band of careful workers, are striking examples of what can be accomplished. Whatever errors future enlightenment may show in their conclusions their ascertained facts will remain positive knowledge; theirs is not what Herr Herman himself described as "pretended authority."

In order to grasp the problems of migration it is necessary to get rid of the puerile and insular aspect of the subject, namely that migrants are merely those birds which come to us, like the swallow and cuckoo in the spring, and those, like the fieldfare and brambling, which visit us in winter but are not with us in summer. The complication of the subject may be demonstrated by a rough classification of the migrants to be observed in the British Islands.

Arbitrary grouping of the members of an avifauna is only for general convenience; many species are represented in more than one group.

When Mr Eagle Clarke was on the Kentish Knock Lightship, off the mouth of the Thames, he found that in autumn there were continuing practically simultaneously the following streams of migration. Immigration from the Continent to England from east to west, and from south-east to north-west, and passage along both lines; emigration from north to south-south-west, and from north-west to south-east, with passage from north to south-south-west. Birds of the same species actually crossed paths, travelling in contrary directions .

The above grouping applies to the British avifauna, but a somewhat similar arrangement might be made of the birds of any particular area, large or small. The grouping of birds for the study of Geographical Distribution is of little consequence in connection with migration, but the mapping of the world into various ornithological rather than zoogeographical regions is of considerable importance, both for convenience in tracing the ranges of migrants, and in the discussion of the history of migration, which almost certainly began in the form of short wanderings from the centres of distribution. It is of comparatively small importance what boundaries we take for the various regions; these depend largely upon the view of certain ornithologists as to which groups of birds shall be considered as typical of the regions in question. Sclater's six regions are perhaps the most universally used. They are as follows:--

Newton suggested an alteration, a continuous northern region to be called the Holarctic Region, which embraces almost the whole of the Northern Hemisphere, and the division of the Australian into Australian and New Zealand Regions. Each of these southern regions is the winter home of some of the Holarctic birds, and it is a matter of dispute whether many of these originated in the northern or southern hemispheres. The value of these artificial divisions of the world is rather in the consideration of the conditions their varied climates and physical features present as attractions to birds in search of suitable nesting places and food supplies.

The study of Migration involves reference to the work of ornithologists of the past and present, the mass of contradictory literature already referred to, and we are repeatedly faced with the difficulty that some particular theory about the vexed questions of the cause or origin of migration, the height and speed at which birds travel, whether they do or do not follow routes, how they find their way, in what order they migrate, how and why they do or do not avoid dangers, or any similar problem, which seems to give finality so far as certain cases are concerned, is met by an absolute negation in other instances. The truth seems clear; more than one factor has influence on most birds, and different species in different places are influenced by different factors. Elliott Coues' sweeping statement, though I strongly disagree with the article in which it occurs, expresses much that is true. "Isepipteses and magnetic meridians, coast-lines and river channels, food-supply and sex-impulses, hunger and love, homing instincts and inherited or acquired memory, thermometer, barometer and hygrometer, may all be factors in the problem, good as far as they function; but none of them, and not all such together, can satisfy the whole equation."

Some of the theses may be laws or rules, but there are no rules without exceptions, and these exceptions may become local rules. Laws regulating migration in one area, whether it be the great continent of America, the British Islands or the islet of Heligoland, may have little application in other parts of the world: local evidence alone can never solve the great problems.

CAUSE AND ORIGIN OF MIGRATION

The question--What makes Birds Migrate? or what causes them to remove from one zone to another at certain seasons, has been answered, no doubt to the satisfaction of the respondents, in many varied ways. Closely connected with the question of immediate impulse is the deeper, and less easy to prove problem as to how migration originated.

It has been dogmatically asserted repeatedly that birds invariably breed in the most northerly part of their range, and winter in the most southerly. Winter, when speaking of Holarctic birds, only applies to the season in the northern hemisphere; the birds which pass south of the equator winter in summer. Whilst accepting this as a rule, two reservations must be made. First, that it only applies to birds of the northern hemisphere, and secondly that it is a rule with exceptions. It seems probable that the breeding area of some of the birds which reach the British Islands in autumn by the so-called east and west route is in more southerly latitudes than our islands, and certainly it seems evident that the temperature of the winter refuge has more effect upon the birds than its geographical position. Perhaps the statement that a bird always nests in the coldest part of its range is more universally correct. Even this may not be invariably the habit, but in acknowledging it as a rule we must clearly understand that this cold district is resorted to at the period of the year when its temperature is at its highest. There are certain birds which breed in Australia and winter in Oceanic islands where the temperature is cooler than in their breeding area.

When considering the migration of birds which summer in the extreme north or breed in the extreme south--alas, but little is known about the migratory habits of many southern breeders--it is comparatively simple to offer an explanation; in the long winter months this home, so desirable in the short weeks of daylight, is dark, ice-bound, and foodless; it is wholly unsuited to the requirements of birds, which, in spite of many assertions to the contrary, have never been proved to hibernate, the only way in which animals can survive for any lengthened period when food supply is entirely cut off.

Birds are structurally provided with the means of escaping from the disastrous effects of adverse circumstances; the power of flight, though not the only way in which animals can migrate, is at the root of the migration of birds. The advantages of the power of flight, to which also it owes its development, include the ability to avoid active and passive enemies, and to remove from one feeding ground to another undeterred by the barriers which restrict the terrestrial animal. A natural sequence of this ability to take advantage of aerial locomotion is the habit of wandering in search of food, more or less noticeable in all birds. The habit of wandering led to the discovery of feeding grounds and suitable nesting places; where these nesting places, probably at first, only removed a short distance from the parents' nesting site, were suitable, dispersal and an extension of the distributional area or range of the species followed; but where the feeding area was unsuited or not so well suited to the needs of the species, hereditary attachment to the original home and memory of the direction of this home, or even in some cases accidental wandering back to the more suitable locality, would originate a migration. Coupled with this are two important factors which would tend to make the habit periodical and regular both as regards time and locality. The memory of the bird, call it instinctive memory if we like, would limit the wanderings in search of food to a certain number of places where food was most abundantly found, and the passage between feeding area and breeding area become regular journeys, at the seasons of the year when an increasing number of young birds in the breeding area drove the overgrown population to seek food further from the base, and again when the sexual impulses urged the birds to seek secure nesting sites. The other factor is the weeding-out influence of mistaken effort, the natural selection which leads to the survival of the fittest. The young wanderer which reached unsuitable lands must either wander further or perish. Judging by the juvenile mortality amongst young birds the failures would be many, and only the successful competitors would return to leave progeny.

Great stress has been laid on the attachment of birds to certain nesting sites, an undoubted fact, and it has been argued that because, in some cases, for hundreds of years certain sites have been occupied by the same species, it is evident that after the death of parents the young will return to and occupy the home. This has even been put forward as evidence that birds do not wander in search of fresh nesting sites. The argument is not sound. It is improbable that in most cases both parents perish in the same year. Birds of prey, and many of the cited instances of long tenancy refer to raptorial birds, have a wonderful power of finding a mate, male or female, to complete the hatching and rearing of the young, when one of a pair has been destroyed. The survivor of any pair might have the home attachment and by bringing a fresh mate create an attachment which would be passed on from mate to mate indefinitely. Again it must not be overlooked that certain sites present advantages to particular species which must be evident to all in search of those advantages; it by no means follows that the occupiers of a nesting site are in any way related, except specifically, to those which occupied it in previous years.

The answer to the argument that birds do not seek fresh nesting places and thus extend their distributional area, is evident when we consider those species which, at the present time, are extending their range. Within the last few years, for instance, the turtle dove and tufted duck have begun to nest regularly in many parts of England in which they were entirely unknown twenty or thirty years ago. The starling has spread and in some parts is spreading still, and many other similar cases might be cited.

In this manner migration, as we know it to-day, may have originated, and as Mr P. A. Taverner expressed it, "however instinctive their habit may now be, there must have been a time when migrations were intelligent movements, intended to escape some danger or secure some advantage" . Granting this, however, as the first cause, we are only on the threshold; the question still remains unanswered, what actually impels the birds to seek fresh food supplies or to look for safe nesting places? The natural answer, the cravings of nature and sexual impulses fails to give satisfaction in every case. Wanderings in search of food might lead in any direction, and probably did in the first place, but now birds in the main travel south in search of food and north in search of home, and many of them perform immense journeys, passing over or through lands which are capable of supporting a wealth of bird-life even in the winter months.

The majority of Arctic birds or those nesting in high latitudes leave before the great harvest of autumn fruits, and even our common swift begins to depart--for all do not go at once--towards the end of July, when insects are more abundant than at any other time of the year. Food supply has not failed when most birds start their journey in search of food! Again in spring, when it is claimed that the powerful sexual impulses are sufficient reason to account for the northward journey, hosts of sexually immature birds and of others which are apparently mature but do not breed that spring, migrate northwards, some even arriving before the mature birds of their own species.

The earlier students of migration insisted that temperature was the sole cause of change of abode; that the northern lands became unsuitable through their falling temperature, and that the birds deserted them for warmer climes, returning when the lands they wintered in became too hot. As a variant of this notion, which cannot be lightly cast aside, the suggestion was mooted that it was not cold but the lack of food during the cold months which drove them south, and that in the Tropics, where at one time it was thought that all migratory birds wintered, food was scarce during the months of extreme heat. Dr. Wallace went further and stated that the incentive to northern migration was the inability to find sufficient soft bodied insects suitable for the nestlings in the Tropics during summer . Yet there are birds which do find food enough for their young, and some of them are insect eaters.

The suggestion that migration owes its origin to the Glacial Epoch, "that supposed solution of so many difficulties," to quote Mr Gadow , has had many exponents. Some take for granted that the Polar Regions were the original home, the centre of dispersal, of all northern birds, and consequently that migration originated in the gradual pushing back of avian life as the ice gained more and more land each year. During the summer, the birds, urged by an irresistible love of home, travelled as far north as the ice allowed them, but gradually they were driven to nest further and further south until they found refuge in the unglaciated parts of the earth. The individuals and the species, if not the whole families of birds, which failed to retreat, went the way of the "thousand types." On the retreat of the ice, the birds, impelled by a mysterious hereditary memory of home and of the good times enjoyed by their remote ancestors, for very very many generations must have been born under more or less sedentary conditions during the Ice Age, began the same pushing forward each year to the limits allowed them. In this case they travelled nearer and nearer to the original home instead of constantly being driven further from it.

Surely the question of original home, at any rate of the home in pre-Glacial days, may be entirely left out of the question. No one can ever prove that this wonderful memory did or could exist. Post-Glacial dispersal northwards, and the foundation of migratory habits of advancing to the new food-producing areas, suitable also for the rearing of young, was doubtless a fact, but would have taken place in any case. The congestion due to the increased numbers driven to a restricted area, would involve a rebound outwards, and the uninhabited areas northward of the refuge would be the natural bourn towards which the birds would travel. The seasonal return of cold would drive them southwards in winter, and the periodical migration habit would thus be originated.

The intense love of home during the spread of glacial conditions would tend rather towards extinction than the formation of any new habits. The birds which possessed the greatest attachment to the particular district would be less likely to fly from adverse conditions, and the reduction of their numbers through the ordinary physiological changes in habit--reduction of the number of young produced, and possibly disinclination to pair--would inevitably end in extinction. The stronger the attachment to home the more likely the bird to remain to the bitter end, and if driven away by increasingly severe winters, to return and attempt to nest in the locality which had become unsuitable for nesting. The spread of glaciation would be gradual and so would be the annihilation of the species, but the end would be sure.

Birds which are cited as species which have shown this remarkable attachment to home, have disappeared before adverse circumstances--the great auk and the Labrador duck.

From what little we do know about the behaviour of our summer birds in their winter home, we may safely conclude that their habits are similar to those of winter visitors to Britain. Only in a few species are there two restricted areas, two abiding places or homes. The necessity of retaining a secure home for the young and the care of these young during their more helpless age keeps the individual birds within a certain area during the breeding season, but at all other times the bird is more or less of a wanderer. The variation, however, of the wanderings is remarkable. For instance the flocks of fieldfares, redwings, and some of the finches which come to winter in the British Islands wander continually from feeding ground to feeding ground, remaining in one place only so long as the food supply is plentiful. When there is a plentiful harvest of beech-mast, chaffinches and bramblings will linger near one clump or avenue of beeches for many weeks, but when, as often happens, the mast crop fails, they become nomadic, and pass from place to place in their hunt for food. They visit fields top-dressed with manure, glean the refuse of the harvest, frequent the farm-yards, and in early spring, visit the budding larches to prey upon their insect pests. On the other hand golden plovers and lapwings are remarkably local in their winter habits, and so long as the weather remains open will frequent the same fields throughout the winter. Severe weather, especially snow, which effectually closes their chance of obtaining food, at once drives them away. They will migrate to the unfrozen mud-flats of the coast, or to those parts of England, generally the south-west, and Ireland, where the climate is normally milder, or they will even leave our islands altogether under great stress.

Any observer may verify the assertion that birds regularly visit certain favourable food-bases by paying attention to the occurrences of birds of passage. The study of a county, for instance, shows that certain species show partiality for particular localities. Thus in Cheshire goldeneyes pass through every spring and autumn, and may be met with occasionally on any of the meres; but at Oakmere, in the Delamere district, one may be almost certain of seeing parties of this species any time during the periods of passage. The curlew may be heard or seen passing over any part of the county, but only in the Delamere fields do we frequently meet with flocks feeding in inland Cheshire. Before the winter resident golden plovers have arrived in autumn and after they have departed in spring, the favourite fields are regularly visited by passing flocks, and the lower reaches of the Mersey, where the common sandpiper is rare as a summer resident, are visited every autumn by parties of birds on passage. Chance may lead a casual wanderer to a good food-supplying spot, but the regularity of appearance suggests habit and memory.

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