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Words: 147150 in 27 pages

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Introduction 531 Acknowledgments 531 Systematic Relationships and Distribution 532 Fossils 534 Economic Importance 534 Study Areas 535 Materials and Methods 537 Terminology 539

Habitat and Limiting Factors 539

Habitat in Kansas 542

Reproduction 543

Mating 543 Insemination 545 Sexual Cycle of Males 545 Sexual Cycle of Females 549 Nesting 554 Eggs 558 Embryonic Development 560 Fertility and Prenatal Mortality 564 Reproductive Potential 565 Number of Reproductive Years 565

Growth and Development 565 Initiation of Growth 565 Size and Appearance at Hatching 566 Growth of Epidermal Laminae 568 Growth of Juveniles 575 Growth in Later Life 578 Annual Period of Growth 580 Environmental Factors Influencing Growth 580 Number of Growing Years 584 Longevity 585 Weight 586 Bony Shell 586 Color and Markings 593 Wear 595

Sexual Dimorphism 595

Temperature Relationships 598 Optimum Temperature 599 Basking 600 Toleration of Thermal Maxima and Minima 601

Hibernation 611

Diet 617

Populations 623

Movements 626 Locomotion 627 Daily Cycle of Activity 629 Seasonal Cycle of Activity 630 Home Range 632 Homing Behavior 636 Social Relationships 637


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